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β-catenin is a major tyrosine-phosphorylated protein during mouse oocyte maturation and preimplantation development

✍ Scribed by Mami Ohsugi; Stefan Butz; Rolf Kemler


Publisher
John Wiley and Sons
Year
1999
Tongue
English
Weight
528 KB
Volume
216
Category
Article
ISSN
1058-8388

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✦ Synopsis


During mouse preimplantation development, the components of the E-cadherincatenin complex are derived from both maternal and zygotic gene activity and the adhesion complex is increasingly accumulated and stored in a nonfunctional form, ready to be used for compaction and the formation of the trophectoderm cell layer (Ohsugi et al., Dev. Dyn. 206:391-402, 1996).

Here, we show that ␤-catenin is a major tyrosine-phosphorylated protein in oocytes and early cleavage-stage embryos and that the relative amount of phosphorylated ␤-catenin is greatly reduced during the morula-blastocyst transition. Peptide-specific antibodies indicate that ␤-catenin undergoes conformational changes and/or that the carboxy-terminal region of ␤-catenin is blocked during preimplantation development. Moreover, the availability of a carboxy-terminal epitope seems to depend on the tyrosine phosphorylation state of ␤-catenin and becomes unmasked when oocytes are treated with the tyrosine kinase inhibitor genistein. Our results suggest that tyrosine phosphorylation of ␤-catenin represents a molecular mechanism to keep the accumulating E-cadherin adhesion complex in a nonfunctional form.