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Structural transition from dimeric to tetrameric i-motif, caused by the presence of TAA at the 3′-end of human telomeric C-rich sequence

✍ Scribed by Mahima Kaushik; Manoj Prasad; Shikha Kaushik; Anju Singh; Shrikant Kukreti


Publisher
Wiley (John Wiley & Sons)
Year
2010
Tongue
English
Weight
833 KB
Volume
93
Category
Article
ISSN
0006-3525

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✦ Synopsis


Abstract

Widely dispersed in genomic DNA, the tandem C‐rich repetitive stretches may fold below physiological pH, into i‐motif structures, stabilized by C·C^+^ pairing. Herein, structural status of a 9‐mer stretch d(CCCTAACCC), [the truncated double repeat of human telomeric sequence], and its extended version, comprising of additional TAA segment at the 3′‐end, representing the complete double repeat d(CCCTAACCCTAA), has been investigated. The pH dependent monophasic UV‐melting, Gel and CD data suggested that while the truncated version adopts a bimolecular i‐motif structure, its complete double repeat (12‐mer) sequence exists in two (bimolecular and tetramolecular) forms. A model is proposed for the tetramolecular i‐motif with conventional C**·C^+^ base pairs, additionally stabilized by asymmetric A·A base pairs at the −3′ TAA flanking ends and Watson–Crick A·**T hydrogen bonding between intervening bases on antiparallel strands. Expanding the known topologies of DNA i‐motifs, such atypical geometries of i‐motifs may have implications in their recognition by proteins. © 2009 Wiley Periodicals, Inc. Biopolymers 93: 150–160, 2010.

This article was originally published online as an accepted preprint. The “Published Online” date corresponds to the preprint version. You can request a copy of the preprint by emailing the Biopolymers editorial office at [email protected]