Latitudinal gradients of tree species composition along the Sierran/Cascade axis in northern California were explored by comparing forests of Lassen Volcanic and Yosemite National Parks, USA. A calibration procedure based on canonical correspondence analysis predicted a mean rate of elevational displacement of 172.1 rn/°latitude for Lassen sites in Yosemite. This is a steep latitudinal gradient compared with other temperate uplands (which average around 100 m/°latitude), but it corresponds with the magnitude of the July mean temperature gradient (! 43 m/°latitude) and the annual precipitation gradient (230 m/°latitude). Elevational displacement of basal-area weighted species means showed considerable variation. The range for montane species was 20--153 m/°latitude; for subalpine species the range was 142-305 m/°latitude. This disparity is related to differential temperature lapse rates between regions and is reinforced by contrasting biogeographic affinities of montane vs. subalpine species. Whereas it is uniformly hot and dry during the growing season at lower elevations in both regions, growing seasons in the subalpine zone are significantly warmer and drier (at comparable elevations) in Yosemite, the more southerly locale. Furthermore, montane species are principally of Sierran affinity, whereas subalpine species are primarily of Pacific Northwestern affinity.