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Kinetic properties of the periplasmic hydrogenase from Desulfovibrio desulfuricans NCIMB 8372 and use in photosensitized H2-production

✍ Scribed by Lars H. Eng; Mona B.-M. Lewin; Halina Y. Neujahr


Publisher
Wiley (John Wiley & Sons)
Year
2007
Tongue
English
Weight
711 KB
Volume
56
Category
Article
ISSN
0268-2575

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✦ Synopsis


Abstract

The periplasmic hydrogenase (hydrogen: ferricytochrome‐c~3~ oxido‐reductase EC 1.12.2.1) from Desulfovibrio desulfuricans (NCIMB 8372) was isolated and the catalytic properties of the enzyme determined with different electron mediators.

In the direction of H~2~‐oxidation with the physiological electron mediator cytochrome c~3~, the value of the apparent second order rate constant (k~cat~/K~m~) is close to that expected for a diffusion‐controlled reaction. In H~2~‐production however, the value of k~cat~/K~m~ with cytochrome c~3~ is almost 200 times lower. When methyl viologen is used as mediator, the value of k~cat~/K~m~ is the same in both directions and is intermediate to the values obtained with cytochrome c~3~.

The close to optimum value of k~cat~/K~m~ in H~2~‐oxidation with cytochrome c~3~ as electron acceptor is discussed with respect to the biological role of this and related hydrogenases.

H~2~‐production occurred with the hydrogenase in a system where the mediator was reduced with either of three photosensitizers (proflavin, 5‐deazariboflavin, Zn‐tetraphenyl porphyrin sulphonate) upon illumination. The efficiency of different mediators is discussed.