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Hyperossification in miniaturized toadlets of the genus Brachycephalus (Amphibia: Anura: Brachycephalidae): Microscopic structure and macroscopic patterns of variation

✍ Scribed by Rute B.G. Clemente-Carvalho; Marta M. Antoniazzi; Carlos Jared; Célio F.B. Haddad; Ana C.R. Alves; Henrique S. Rocha; Gabriela R. Pereira; Davi F. Oliveira; Ricardo T. Lopes; Sergio F. dos Reis


Book ID
102904818
Publisher
John Wiley and Sons
Year
2009
Tongue
English
Weight
708 KB
Volume
270
Category
Article
ISSN
0362-2525

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✦ Synopsis


Abstract

Species of the genus Brachycephalus, have a snout‐vent length of less than 18 mm and are believed to have evolved through miniaturization. Brachycephalus
ephippium, is particularly interesting; because its entire skull is hyperossified, and the presacral vertebrae and transverse processes are covered by a dorsal shield. We demonstrate in this paper that, at the macroscopic level, a completely hyperossified skull and dorsal shield occur only in B. ephippium, but not in B. ferruginus, B. izechsohni, B. pernix, B. pombali, B. brunneus, B. didactylus, and B. hermogenesi. An intermediate condition, in which the skull is hyperossified but a dorsal shield is absent, occurs in B. vertebralis, B. nodoterga, B. pitanga, and B. alipioi. The microscopic structure of hyperossification was examined in skulls of B. ephippium and B. pitanga, revealing a complex organization involving the presence of Sharpey fibers, which in humans are characteristic of periodontal connections. J. Morphol., 2009. © 2009 Wiley‐Liss, Inc.