Global changes in amide hydrogen exchange rates for a protein antigen in complex with three different antibodies : David C. Williams Jr.a, David C. Benjamina, Roberto J. Poljakd, Gordon S. Rulebc. aDepartment of Microbiology, bDepartment of Biochemistry, cthe Beirne B. Carter Center for Immunological Research., University of Virginia, Charlottesville, VA 22908, dCenter for Advanced Research in Biotechnology, University of Maryland Biotechnology Institute, 9600 Gudelsky Drive, Rockville, Maryland 20850, USA