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Biochemical analysis of mutants defective in nitrate assimilation inNeurospora crassa: Evidence for autogenous control by nitrate reductase

✍ Scribed by A. Brian Tomsett; Reginald H. Garett


Publisher
Springer
Year
1981
Tongue
English
Weight
849 KB
Volume
184
Category
Article
ISSN
1617-4615

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✦ Synopsis


A biochemical analysis of mutants altered for nitrate assimilation in Neurospora crassa is described. Mutant alleles at each of the nine nit (nitrate-nonutilizing) loci were assayed for nitrite reductase activity, for three partial activities of nitrate reductase, and for nitrite reductase activity. In each case, the enzyme deficiency was consistent with data obtained from growth tests and complementation tests in previous studies. The mutant strains at these nit loci were also examined for altered regulation of enzyme synthesis. Such experiments revealed that mutations which affect the structural integrity of the native nitrate reductase molecule can result in constitutive synthesis of this enzyme protein and of nitrite reductase. These results provide very strong evidence that, as in Aspergillus nidulans, nitrate reductase autogenously regulates the pathway of nitrate assimilation. However, only mutants at the nit-2 locus affect the regulation of this pathway by nitrogen metabolite repression.