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A disulfide bridge near the active site of carbapenem-hydrolyzing class A β-lactamases might explain their unusual substrate profile

✍ Scribed by Xavier Raquet; Josette Lamotte-Brasseur; Fabrice Bouillenne; Jean-Marie Frère


Publisher
John Wiley and Sons
Year
1997
Tongue
English
Weight
306 KB
Volume
27
Category
Article
ISSN
0887-3585

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✦ Synopsis


Bacterial resistance to ␤-lactam antibiotics, a clinically worrying and recurrent problem, is often due to the production of ␤-lactamases, enzymes that efficiently hydrolyze the amide bond of the ␤-lactam nucleus. Imipenem and other carbapenems escape the activity of most active site serine ␤-lactamases and have therefore become very popular drugs for antibacterial chemotherapy in the hospital environment. Their usefulness is, however, threatened by the appearance of new ␤-lactamases that efficiently hydrolyze them. This study is focused on the structure and properties of two recently described class A carbapenemases, produced by Serratia marcescens and Enterobacter cloacae strains and leads to a better understanding of the specificity of ␤-lactamases. In turn, this will contribute to the design of better antibacterial drugs. Threedimensional models of the two class A carbapenemases were constructed by homology modeling. They suggested the presence, near the active site of the enzymes, of a disulfide bridge (C69-C238) whose existence was experimentally confirmed. Kinetic parameters were measured with the purified Sme-1 carbapenemase, and an attempt was made to explain its specific substrate profile by analyzing the structures of minimized Henri-Michaelis complexes and comparing them to those obtained for the ''classical'' TEM-1 ␤-lactamase. The peculiar substrate profile of the carbapenemases appears to be strongly correlated with the presence of the disulfide bridge between C69 and C238.